The California Department of Public Health and Environment has issued $1 million in fines to the UC Berkeley Botanical Garden for violations stemming from an April 2015 outbreak of coronavirus, the UC Santa Cruz Department of Animal and Plant Health has said.

The fines are the latest in a string of financial penalties levied by the university over the outbreak, according to a UCSC statement.

The university has paid $2.1 million and is working to reduce the cost of the pandemic-related fines, the statement said.

In April, the university announced that it had imposed a $2 million financial penalty against the Botanical Gardens for improper animal use.

In June, the botanical campus was fined $4.2 million by the state for improper use of animals and plants.

UC Berkeley also has $4 million in veterinary fines and is seeking to reduce costs, the department said.

More on botanical Gardens:UC Berkeley Botanic Gardens fined $1,000,000 for improper animals use and $4,000K for veterinary violations.

UCSC is working on reducing costs and reducing the cost.